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Paleontology or palaeontology (/ˌplɪɒnˈtɒləi/, /ˌplɪənˈtɒləi/ or /ˌpælɪɒnˈtɒləi/, /ˌpælɪənˈtɒləi/) is the scientific study of prehistoric life. It includes the study of fossils to determine organisms' evolution and interactions with each other and their environments (their paleoecology). As a "historical science" it attempts to explain causes rather than conduct experiments to observe effects. Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term itself originates from Greek: παλαιός (palaios) meaning "old, ancient," ὄν, ὀντ- (on, ont-), meaning "being, creature" and λόγος (logos), meaning "speech, thought, study". Paleontology lies on the border between biology and geology. It now uses techniques drawn from a wide range of sciences, including biochemistry, mathematics and engineering. Use of all these techniques has enabled paleontologists to discover much of the evolutionary history of life, almost all the way back to when Earth became capable of supporting life, about 3,800 million years ago. As knowledge has increased, paleontology has developed specialized sub-divisions, some of which focus on different types of fossil organisms while others study ecology and environmental history, such as ancient climates. Body fossils and trace fossils are the principal types of evidence about ancient life, and geochemical evidence has helped to decipher the evolution of life before there were organisms large enough to leave fossils. (see more...)




Illustration of the skull of Edaphosaurus in lateral and dorsal view from the book "Water Reptiles of the Past and Present" by Samuel Wendell Williston.

The history of paleontology traces the history of the effort to study the fossil record left behind by ancient life forms. Although fossils had been studied by scholars since ancient times, the nature of fossils and their relationship to life in the past became better understood during the 17th and 18th centuries. At the end of the 18th century the work of Georges Cuvier ended a long running debate about the reality of extinction and led to the emergence of paleontology as a scientific discipline.

The first half of the 19th century saw paleontological activity become increasingly well organized. This contributed to a rapid increase in knowledge about the history of life on Earth, and progress towards definition of the geologic time scale. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. After Charles Darwin published Origin of Species in 1859, much of the focus of paleontology shifted to understanding evolutionary paths.

The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection, as demonstrated by a series of important discoveries in China near the end of the 20th century. There was also a renewed interest in the Cambrian explosion that saw the development of the body plans of most animal phyla. (see more...)





Photograph of Edward Drinker Cope

Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Cope distinguished himself as a child prodigy, publishing his first scientific paper at the age of nineteen. Cope later married and moved from Philadelphia to Haddonfield, New Jersey, although Cope would maintain a residence and museum in Philadelphia in his later years.

Cope had little formal scientific training, and he eschewed a teaching position for field work. He made regular trips to the American West prospecting in the 1870s and 1880s, often as a member of United States Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition now known as the Bone Wars. Cope's financial fortunes soured after failed mining ventures in the 1880s. He experienced a resurgence in his career toward the end of his life before dying in 1897.

Cope's scientific pursuits nearly bankrupted him, but his contributions helped to define the field of American paleontology. He was a prodigious writer, with 1,400 papers published over his lifetime, although his rivals would debate the accuracy of his rapidly published works. He discovered, described, and named more than 1,000 vertebrate species including hundreds of fishes and dozens of dinosaurs. His proposals on the origin of mammalian molars and for the gradual enlargement of mammalian species over geologic time ("Cope's Law") are notable among his theoretical contributions. (see more...)





Edward Drinker Cope (left) and Othniel Charles Marsh (right).

The Bone Wars is the name given to a period of intense fossil speculation and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope and Othniel Charles Marsh. The two paleontologists used underhanded methods to out-compete the other in the field, resorting to bribery, theft, and destruction of bones. The scientists also attacked each other in scientific publications, attempting to ruin the other's credibility and cut off his funding.

Originally colleagues who were civil to each other, Cope and Marsh became bitter enemies after several personal slights between them. Their pursuit of bones led them west to rich bone beds in Colorado, Nebraska, and Wyoming. From 1877 to 1892, both paleontologists used their wealth and influence to finance their own expeditions and to procure services and fossils from dinosaur hunters. By the end of the Bone Wars, both men exhausted their funds in fueling their intense rivalry.

Cope and Marsh were financially and socially ruined by their efforts to disgrace each other, but their contributions to science and the field of paleontology were massive; the scientists left behind tons of unopened boxes of fossils on their deaths. The feud between the two men led to over 142 new species of dinosaurs being discovered and described. Several historical books and fictional adaptations have also been published about this period of intense paleontological activity. (see more...)





Edward Drinker Cope (left) and Othniel Charles Marsh (right).

Bone Sharps, Cowboys, and Thunder Lizards: A Tale of Edward Drinker Cope, Othniel Charles Marsh, and the Gilded Age of Paleontology (2005) is a graphic novel written by Jim Ottaviani and illustrated by the company Big Time Attic. The book tells a slightly fictionalized account of the Bone Wars, a period of intense excavation, speculation, and rivalry which led to a greater understanding of dinosaurs in the western United States. This novel is the first semi-fictional work written by Ottaviani; previously, he had taken no creative license with the characters he depicted, portraying them strictly according to historical sources. Bone Sharps follows the two scientists Edward Drinker Cope and Othniel Marsh as they engage in an intense rivalry for prestige. Ottaviani has Cope and Marsh interact and meet many important figures of the Gilded Age, from P. T. Barnum to U.S. Grant, as the two scientists pursue their hotheaded and sometimes illegal acquisitions of fossils. Unlike in his previous books, "the scientists are the bad guys this time". Upon release, the novel received praise from critics for its exceptional historical content, although some reviewers wished more fiction had been woven into the story. (see more...)





The Tree of Life as depicted by Ernst Haeckel in The Evolution of Man (1879) illustrates the 19th-century view that evolution was a progressive process leading towards man.

Evolutionary thought, the conception that species change over time, has roots in antiquity. With the beginnings of biological taxonomy in the late 17th century, a new anti-Aristotelian approach to modern science challenged traditional essentialism. Naturalists began to focus on the variability of species; the emergence of paleontology with the concept of extinction further undermined the static view of nature. In the early 19th century, Jean-Baptiste Lamarck proposed the first fully formed theory of evolution.

In 1858, Charles Darwin and Alfred Russel Wallace published a new evolutionary theory that was explained in detail in Darwin's On the Origin of Species (1859). Unlike Lamarck, Darwin proposedcommon descent and a branching tree of life. The theory was based on the idea of natural selection, and it synthesized a broad range of evidence from animal husbandry, biogeography, geology, morphology, and embryology.

The debate over Darwin's work led to the rapid acceptance of the general concept of evolution, but the specific mechanism he proposed, natural selection, was not widely accepted until it was revived by developments in biology that occurred during the 1920s through the 1940s. Before that time most biologists argued that other factors were responsible for evolution. The synthesis of natural selection with Mendelian genetics during the 1920s and 1930s founded the new discipline of population genetics. Throughout the 1930s and 1940s, population genetics became integrated with other biological fields, resulting in a widely applicable theory of evolution that encompassed much of biology—the modern evolutionary synthesis. (see more...)